He began his graduate work with Professor Rychnovsky at the University of Minnesota in 1991 and moved with him to UCI, where he received his Ph.D. degree in 1997. Currently, he is an NIH postdoctoral fellow at UCI with Larry Overman. Timothy Richardson received his B.S. degree in biochemistry from the College of Biological Sciences at the University of Minnesota in 1993. He received a Ph.D. in Chemistry in 1997 from the University of Minnesota and is currently working as an NIH postdoctoral fellow in the laboratories of David Evans at Harvard University.