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Invasive plant species have been hypothesized to possess a suite of traits associated with successful invasion into areas in which they are introduced (van Kleunen et al., 2010;Rejmánek & Richardson, 1996).For instance, invasive species often have functional traits that confer greater resource capture ability, such as larger specific leaf areas (SLA) and smaller root-to-shoot ratios (R/S ratios) (Ehrenfeld, 2003;Grotkopp & Rejmánek, 2007). These traits result in faster growth rates and higher reproductive outputs of invasive species than those

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