Fossil associations from the middle and upper Eocene (Bartonian and Priabonian) sedimentary succession of the Pamplona Basin are described. This succession was accumulated in the western part of the South Pyrenean peripheral foreland basin and extends from deep-marine turbiditic (Ezkaba Sandstone Formation) to deltaic (Pamplona Marl, Ardanatz Sandstone and Ilundain Marl formations) and marginal marine deposits (Gendulain Formation). The micropalaeontological content is high. It is dominated by foraminifera, and common ostracods and other microfossils are also present. The fossil ichnoasssemblages include at least 23 ichnogenera and 28 ichnospecies indicative of Nereites, Cruziana, and ?Scoyenia-Mermia ichnofacies. Body macrofossils of about 80 taxa corresponding to macroforaminifera, sponges, corals, bryozoans, brachiopods, annelids, molluscs, arthropods, echinoderms and vertebrates have been identified. Both the number of ichnotaxa and of species (e. g. bryozoans, molluscs and condrichthyans) may be considerably higher. Body fossil assemblages are comparable to those from the Eocene of the Nord Pyrenean area (Basque Coast), and also to those from the Eocene of the west-central and eastern part of South Pyrenean area (Aragon and Catalonia). At the European scale, the molluscs assemblages seem endemic from the Pyrenean area, although several Tethyan (Italy and Alps) and Northern elements (Paris Basin and Normandy) have been recorded. Palaeontological data of studied sedimentary units fit well with the shallowing process that throughout the middle and late Eocene occurs in the area, according to the sedimentological and stratigraphical data.Se describen las asociaciones fósiles del Eoceno medio y superior (Bartoniense y Priaboniense) de la sucesión sedimentaria de la Cuenca de Pamplona. Esta sucesión se acumuló en la parte oeste de la Cuenca de antepaís periférica surpirenaica y comprende desde turbiditas marinas profundas (Formación Areniscas de Ezkaba) hasta depósitos deltaicos (Formaciones Margas de Pamplona, Areniscas de Ardanatz y Margas de Ilundain) y marinos marginales (Formación Gendulain). El contenido micropaleontológico es alto y está dominado por los foraminíferos. Los fósiles de ostrácodos y otros organismos son también abundantes. Las asociaciones de icnofósiles incluyen al menos 23 icnogéneros y 28 ichnoespecies, indicativas de las icnofacies de Nereites, Cruziana, y ?Scoyenia-Mermia. Se han identificado fósiles corporales de unos 80 taxones, correspondientes a macroforaminíferos, esponjas, corales, briozoos, braquiópodos, anélidos, moluscos, artrópodos, equinodermos y vertebrados. El número de icnotaxones y de especies (p. e. briozoos, moluscos gasterópodos, bivalvos y peces condríctios) puede ser considerablemente mayor. Las asociaciones de fósiles corporales son comparables a las del Eoceno del Área norpirenaica (Costa vasca) y de las partes central y este del Área surpirenaica (Aragón y Cataluña). A escala europea, la asociación de moluscos parece endémica del Área pirenaica,...

Gastropods and bivalve associations from the middle and ?upper Eocene (Bartonian and ?Priabonian) sedimentary succession of the Pamplona Basin are described. This succession was accumulated in the western part of the South Pyrenean peripheral foreland basin and extends from deep marine turbiditic (Ezkaba Sandstone Formation) to deltaic (Pamplona Marl, Ardanatz Sandstone and Ilundain Marl formations) and marginal marine deposits (Gendulain Formation). Fossils are generally fragmentary and many correspond to remains that have undergone taphonomic reworking. Significant effects of diagenesis are also detected, with evidence of compression or deformation, dissolution, aragonitecalcite neomorfism and cementation by celestite. Bioerosion traces and fossil encrusters are common. Fossils of 37 taxa, 25 gastropods and 12 bivalves, have been identified confidently. The number of species is probably considerably higher, as the study of some fossils, mainly the most small-sized GEODIVERSITAS • 2018 • 40 (11) Astibia H. et al. MOTS CLÉS Gastéropodes, bivalves, Paléogène, sédiments marins peu profonds, Pyrénées, combinaisons nouvelles, espèce nouvelle. species, is pending. One new species, Athleta (Volutospina) delvallei Astibia, Merle & Pacaud, n. sp. (Gastropoda, Volutidae), is described herein. Fossil assemblages are comparable to those from the Eocene of the Basque Coast (North Pyrenean area) and Aragon and Catalonia (west-central and eastern part of South Pyrenean area). Most of the mollusc taxa seem to be endemic to the Pyrenean area, but several Tethyan and Northern elements have also been recorded. These results enlarge the database for a better understanding of the evolution of global marine biodiversity throughout the Eocene. RÉSUMÉ Gastéropodes et bivalves des formations marneuses de l'Éocène du Bassin de Pampelune et de ses environs (Navarre, Pyrénées occidentales). Les associations de gastéropodes et de bivalves de la succession sédimentaire de l'Éocène moyen et ?supérieur (Bartonien et ?Priabonien) du Bassin de Pampelune sont décrites. Cette succession s'est accumulée dans la partie occidentale du bassin d'avant-pays périphérique sud-pyrénéen et comprend des dépôts marins turbiditiques profonds (Formation des Grès d'Ezkaba) des dépôts deltaïques (formations des Marnes de Pampelune, des Grès d'Ardanatz et des Marnes d'Ilundain) et des dépôts marins marginaux (Formation Gendulain). Les fossiles sont généralement fragmentaires et beaucoup d'entre eux correspondent à des restes remaniés. Des effets significatifs de la diagénèse sont également détectés, avec des signes de compression ou de déformation, de dissolution, de néomorphisme aragonite-calcite et de cimentation par célestine. Les traces de bioérosion et les fossiles d'organismes incrustants sont fréquents. Les restes fossiles de 37 espèces, dont 25 correspondant à des gastéropodes et 12 à des bivalves, ont été identifiés. Ce nombre pourrait être considérablement plus élevé, grâce à l'étude de très petites espèces (micro-mollusques) qui reste à faire. Une nouve...

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and Ampullospira) and three subgenera (Globularia, Deshayesia, and Cernina within the genus Globularia). Forty−six Cainozoic species revealed residual colour patterns and 29 of them, belonging to six genera (Ampullina, Globularia, Crommium, Amaurellina, Pachycrommium, Amauropsina)