Th e polydesmoid family Haplodesmidae Cook, 1895 is reviewed and shown to be a senior subjective synonym of the family Doratodesmidae Cook, 1896, syn. n. Th e Haplodesmidae therefore encompasses six genera and 30 recognizable species, all keyed here, including the following six new species: Eutrichodesmus basalis sp. n. and E. armatocaudatus sp. n. from Vietnam; E. communicans sp. n. from Vanuatu, Melanesia; and E. latus sp. n., E. similis sp. n. and E. incisus sp. n. from China. Th e following new synonymies are proposed:

Eutrichodesmus, the largest genus in the Oriental family Haplodesmidae, is reviewed and shown to encompass 24 recognizable species, all keyed, including the following nine new species: E. regularis sp. n., E. aster sp. n., E. fi lisetiger sp. n., E. curticornis sp. n., E. asteroides sp. n. and E. griseus sp. n. from Vietnam, E. distinctus sp. n. from China, E. multilobatus sp. n. from Laos, and E. reductus sp. n. from Sulawesi, Indonesia. Th is genus is slightly redefi ned as follows: Gonopod coxae usually abundantly setose ventrolaterally; telopodite usually slender, not enlarged towards end of femorite, but typically with a more or less distinct process or outgrowth laterally, opposite recurvature point of seminal groove; solenomere thereafter usually comprising most of telopodite, sometimes elaborate; seminal groove normally terminating distally to subapically, with or without a hairpad; acropodite normally small to nearly absent. Golovatch et al. / ZooKeys 12: 1-46 (2009) 2

In addition to one of the two species of Trichopolydesmidae hitherto recorded from Cameroon, Polydesmusintegratus Porat, 1894, which is revised based on type material and shown to represent the genus Hemisphaeroparia Schubart, 1955, comb. n., 12 new species from the same genus are described from that country: H.zamakoesp. n., H.bangoulapsp. n., H.spinigersp. n., H.ongotsp. n., H.digitifersp. n., H.parvasp. n., H.fuscasp. n., H.bonakandasp. n., H.bamboutossp. n., H.subfalcatasp. n., H.falcatasp. n. and H.mouankosp. n. A key to all 13 species (of Hemisphaeroparia) known to occur in Cameroon is presented, and their distributions are mapped. All ten recognizable (but excluding two dubious) Afrotropical genera or subgenera of Trichopolydesmidae are rediagnosed and reclassified, based both on their type species and a presumed scenario of gonopodal evolution. As a result, the number of accepted genera is reduced to five: Sphaeroparia Attems, 1909 (= Megaloparia Brolemann, 1920), Physetoparia Brolemann, 1920 (= Elgonicola Attems, 1939, syn. n., = Mabocus Chamberlin, 1951, syn. n., = Heterosphaeroparia Schubart, 1955, syn. n.}, Eburodesmus Schubart, 1955, Mecistoparia Brolemann, 1926 (= Dendrobrachypus Verhoeff, 1941, syn. n.), and Hemisphaeroparia.

The javanicus-group of the basically southeast Asian genus Glyphiulus is shown to currently comprise 22 species, of which four are new: G. echinoides sp.n., from a cave in Guangxi Province, China, as well as G. subechinoides sp.n., G. parechinoides sp.n. and G. submediator sp.n., all three latter species from Vietnam. Besides this, a new record is provided, and additional abundant illustrations are presented, for G. mediobliteratus Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007, from a cave in northeastern Vietnam. The species G. vietnamicus Mauriès, 1977, from a cave in northern Vietnam, is formally ejected from the javanicus-group, instead being transferred to Hypocambala Silvestri, 1897; this results in H. vietnamica (Mauriès, 1977), comb.n. ÐÅÇÞÌÅ. Ãðóïïà âèäîâ javanicus èñêîííî þãî-âîñòî÷íîàçèàòñêîãî ðîäà Glyphiulus òåïåðü ñîäåðaeèò 22 âèäà, âêëþ÷àÿ ÷åòûðå íîâûe: G. echinoides sp.n. èç ïåùåðû â ïðîâèíöèè Ãóàíñè (þaeíûé Êèòàé), G. subechinoides sp.n., G. parechinoides sp.n. è G. submediator sp.n., âñå òðè ïîñëåäíèõ âèäà èç Âüåòíàìà. Êðîìå òîãî, çàðåãèñòðèðîâàíà è áîãàòî ïðîèëëþñòðèðîâàíà íîâàÿ íàõîäêà G. mediobliteratus Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007 â ïåùåðå íà ñåâåðî-âîñòîêå Âüåòíàìà. Âèä G. vietnamicus Mauriès, 1977 èç ïåùåðû â Ñåâåðíîì Âüåòíàìå ôîðìàëüíî èñêëþ÷åí èç ãðóïïû javanicus è ïåðåâåäåí â ñîñòàâ ðîäà Hypocambala Silvestri, 1897, â ðåçóëüòàòå ÷åãî âîçíèêàåò H. vietnamica (Mauriès, 1977), comb.n.

The basically southeast Asian genus Plusioglyphiulus is shown to currently comprise 27 species, all keyed, of which 13 are new, stemming from Thailand, and mapped: P. panhai sp. n., P. antiquior sp. n., P. sutchariti sp. n., P. likhitrakarni sp. n., P. pimvichaiae sp. n., P. erawan sp. n., P. wat sp. n., P. tham sp. n., P. phra sp. n., P. jaydee sp. n., P. puttakun sp. n., P. saksit sp. n. and P. samakkee sp. n. Although the new species have mainly been collected in caves, none seems to actually represent a troglobiont. A second record of P. ampullifer Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2009 is provided in southern Vietnam, as well as new illustrations are presented for P. bessoni Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2009, prepared from strictly topotypic samples in northern Thailand. With the discovery of P. panhai sp. n. and, especially, P. antiquior sp. n., both from southern Thailand, and both markedly transitional to the more northerly diversified javanicus-group of Glyphiulus, the genus Plusioglyphiulus is reconfirmed as being heterogeneous. Its diagnosis seems to be based now only on a single, rather weak apomorphy in posterior gonopod structure. Some evolutionary and zoogeographical considerations are presented to substantiate the predominantly eastward and/or southward speciation events in typical Plusioglyphiulus which have reached northern and eastern Borneo in the east and southeast.