The core group of Euomphalidae with the genera Euomphalus, Straparollus, Serpulospira, Phymatifer, Schizostoma, Nodeuomphalus n. g. (Devonian to Permian) are characterized by a cyrtoconic, openly coiled, planispiral protoconch. This character distinguishes them from members of the four extant subclasses of the Gastropoda (Archaeogastropoda, Neritimorpha, Caenogastropoda and Heterostropha) and places the Euomphaloidea at a similarly high taxonomic level, the new subclass Euomphalomorpha. With Euomphalopsis the group may have made its first appearanee in the Late Cambrian, and it is certainly present from the Mid Devonian to Late Permian and thus the range is about 300 million years in duration. Throughout the existence of the Euomphaloidea other gastropod units developed convergent shell forros which can be detected by the morphology of the protoconch and, in well preserved fossils, the mineralogical and stmctural composition of the she[l. The new taxa Euomphalomorpha n. subclassis, Nodeuomphalus n, g., N. paffrathianus n. sp., and "Ptanerotinus "" scheeri n. sp. are describe& K e y w o r d s : Gastropoda, Euomphalidae, taxonomy, protoconch, Paleozoic. Kurzfassung[Die systematische Stellung der Euomphalidae (Gastropoda).]-Den Kernbereich der Familie Euomphalidae stellen die Gattungen Euomphalus, Straparollus, Serpulospira, Phymatifer, Schizostoma und Nodeuomphalus aus dem Zeitraum Devon bis Perro. Sie werden durch einen Protoconch gekennzeichnet, der mit einer eif'6rmigen Anfangskalotte ausgestattet ist, an die sich eine erste offene Windung anschlieBt. Dadurch verbleibt eine zentrale Offnung im Inneren der ersten planspiraligen Windung sichtbar im Apex der Schale. Dieser Protoconch unterscheidet die neue Unterklasse Euomphalomorpha von den auch heute noch existenten Unterklassen der Gastropoda, den Archaeogastropoda, Neritimorpha, Heterostropha und Caenogastropoda. Mit Euomphalopsis war m6glicherweise schon ira sp/iten Kambrium ein Vertreter der Euomphalomorpha present. Vertreter der Gruppe sind vom Mitteldevon bis zum spfiten Perm gesichert nachzuweisen. Schon seit dem Ordovizium treten innerhalb anderer Gastropoden-Taxa konvergente Arten auf, so daB nur bei einer Erhaltung des Protoconches, oder auch der Schalenstruktur, Euomphalomorpha gesichert erkannt werden k6nnen. Die neuen Taxa Euomphalomorpha n. subclassis, Nodeuomphalus n. g., AL pafJ?athianus n. sp., and ,, Planerotinus" scheeri n. sp. werden beschrieben.
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