Amanita has traditionally been divided into two subgenera based on basidiospore amyloidity (e.g. Singer 1986). Subgenus Amanita includes those species with inamyloid basidiospores, and has traditionally been subdivided into sections Amanita, Vaginatae, and Caesareae based on macromorphology (e.g. Moser 1967). Molecular studies have corroborated these sections by demonstrating the monophyly of each (Weiss et al. 1998, Tang et al. 2015). Subgenus Lepidella includes those species with amyloid basidiospores and has been further subdivided into four sections based on velar morphology: sect. Amidella (marginal appendiculae and saccate volva present), sect. Lepidella (marginal appendiculae present, saccate volva absent), sect. Phalloideae (marginal appendiculae absent, saccate volva present), and sect. Validae (marginal appendiculae and saccate volva absent) (Corner & Bas 1962). Molecular studies have supported the monophyly of sect. Amidella and sect.

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