Due to their rapid proliferation, persistence and tolerance of disturbance, some creeping perennial herbs have become invasive and harmful weeds in natural and semi-natural ecosystems, as well as in production systems. The grass Elymus repens (L.) Gould. (Couch grass) and the dicotyledonous Sonchus arvensis L.(Perennial sow-thistle) are serious agricultural weeds in the temperate climate zones of the world, especially in organic farming (Ringselle et al., 2020;Salonen et al., 2011). As agricultural weeds, they primarily grow in light-rich environments. The dicotyledonous Aegopodium podagraria L. (Ground elder) is a shade-tolerant

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