The genera Anemonastrum and Knowltonia earlier usually included in Anemone sensu lato (Ranunculaceae: Anemoneae) were recently proposed to be recognized in expanded circumscriptions, based on molecular phylogenetic findings and partly also morphological and biogeographical evidence. Both these genera contain some representatives that are noteworthy from morphological and biogeographical viewpoints, and the general ranges of these genera (combining the ranges of their included species) demonstrate remarkable continent-scale disjunctions. In particular, Knowltonia in its new circumscription (corresponding to Anemone sect. Pulsatilloides sensu Hoot et al., 2012) contains southern African and some American taxa (mostly South American ones, including Oreithales, Barneoudia, etc.), and K. crassifolia from Tasmania. Anemonastrum (corresponding to Anemone subg. Anemonidium sensu Hoot et al., excluding Hepatica) is mainly Eurasian, but it also includes several North American species, as well as A. antucense from southern South America and A. tenuicaule from New Zealand. Species of the newly outlined Anemonastrum and Knowltonia were placed in various infrageneric taxa of Anemone sensu lato. Following the re-circumscription of genera, it is now timely to nomenclaturally update and partly re-circumscribe the relevant infrageneric taxa earlier treated in Anemone sensu lato. Here we propose the following new combinations at the section and subsection ranks: Knowltonia sect.
At the results of investigation of micromorphology features of seed surface of the Portulaca oleracea aggr. (Portulacaceae Juss.) from Chernivtsi Region (Ukraine) six morphotypes of this taxa were recorded. Some peculiarities of the seed morphology of Portulaca L., and features of the ultrastructure of seed surface of the P. oleracea aggr. morphotypes, based on the results of micromorphological studies with using electronic scanning microscope was described. The general for the representative of the genus (e.g. globous, ovate or almost reniform shaped of seeds; black or dark brown color) and specify for the morphological types (star-shaped, isodiametric or elongate cells of seed surface with central part and long or short rays; smooth surface of seed or with knobs (from 0,03 mm in diameter) or papillae (0,01–0,02 mm), etc.) peculiarities are determined. The original scanning electron microscopy (SEM) photographs of seed surface ultrastructure of the studied morphological types and key fragment for its determination are suggested. According to our investigation six morphotypes of Portulaca oleracea aggr. are presented in Chernivtsi Region. Some of them are new for studied territory (P. granulatostellulata (Poelln.) Ricceri & Arrigoni, P. papillatostellulata (Danin & H.G. Baker) Danin), and other (Portulaca nitida (Danin & H.G. Baker) Ricceri & Arrigoni, P. trituberculata Danin, Domina & Raimondo, P. tuberculata (Danin & H.G. Baker) Danin) – new floristic records for Ukraine, and one of them (P. oleracea) – was known before also. The maps of distribution of this morphotypes in the region are presented. At the results of morphotypes distribution the most common is P. granulatostellulata and P. oleracea – rare in studied region like as in Europe also.
The results of studying of seeds of the Portulaca oleracea L. (Portulacaceae Juss.) from Transcarpathia (Ukraine) are presented in the publication. The seed surface ultrastructure of taxa of the P. oleracea-aggr. based on the results of micromorphological studies with using electronic scanning microscope was described. The general for the representative of the genus (e.g. globous, ovate or almost reniform shaped of seeds; dark) and specify for the morphological types (star-shaped, isodiametric or elongate cells of seed surface with central part and long or short rays; smooth surface of seed or with knobs (from 0,03 mm in diameter) or papillae (0,01–0,02 mm in diameter), etc.) peculiarity are determined. The original figures of seed surface ultrastructure of the studied morphological types and key for its determination are prepared. As a results of investigation the new for the Ukrainian flora taxa, P. cypria Danin, P. granulatostellulata (Poelln.) Ricceri & Arrigoni and P. papillatostellulata (Danin & H.G. Baker) Danin, from Transcarpathia are presented. All studied species are: kenophytes according time of immigration; xenophytes, according way of distribution; and ephemerophytes, according degree of naturalization. The map of distribution of the taxa in the region is presented.
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