and BELSPO (Politique scientifique fédérale, Belgium). We were very grateful to Valentin Yamani and Alexane Broussin who kindly collected carabid species. We also thank the farmers for allowing us to sample their hedgerows. We thank LETG-Rennes UMR 6554 CNRS for providing land cover data of the Zone Atelier Armorique (ZAAr). We thank the French LTER (Long Term Clean manuscriptClick here to access/download;Manuscript;Bergerot_R4_clean_version.docx Click here to view linked References