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Authors' Contribution QJ planned the work, did the experimental lab work and wrote the primary manuscript. AS supervised the work. SBR helped in data analysis, MA helped in prepration of the manuscript.

Authors' Contribution QJ and SBR conceived and designed the study and wrote the article. QJ and MI collected the specimens and identified them. SBR and SU made the altitude maps. MA, FZ and QZ helped in acquisition of data and edited the manuscript

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