Recently, clove plantation was mostly established in monoculture systems for production efficiency (Arimalala et al., 2019; Danthu et al., 2014; Martin, 1991). The frequent litter collection associated with intensive soil surface disturbances in a whole area of the crop field, however, is causing deterioration plant species diversity of ground vegetation. Therefore, mixed culture by intercropping of clove with other crop species is suggested as an alternative method which would reduce the impact of frequent litter collection on biodiversity of ground vegetation for sustainable management of clove production. Though the mixed culture is expected to have ecological advantages in terms of biodiversity conservation, at the same time, it will also bring inter-species competition of resources such as light or soil water (

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