Potassium (K + ), the most abundant inorganic cation in plant cells, is essential for plant growth and thus strongly determines crop yield (Adams & Shin, 2014;Wang & Wu, 2017). The regulation of K + in plant health is involved in many aspects of cell physiology and metabolism, including cell expansion, enzyme activation, stomatal opening, and turgor pressure maintenance (Anschütz et al., 2014).Considering the significant roles of K + in crop yield and plant health, improving K + use efficiency in plants is emerging as an important