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Ornamental crabapples (Malus sp.), which are shrubs or small trees in the rose family (Tang et al. 2015), have a long history of cultivation and are well-known as the "national beauty" in China (Pang 2008). The plants are popular ornamentals because of their diversity of habit, including columnar, fastigiate, upright, spreading, drooping, and weeping; various flower colors, such as white, pink, red, and purple; colorful foliage, such as yellow-green, green, gradient, red-brown, purple; and gorgeous autumn and winter fruit