Smk1 is a meiosis-specific MAP kinase that is activated by a binding partner, Ssp2. This study shows that the meiosis-specific Cdc20 homologue, Ama1, triggers Ssp2/Smk1 complex formation at specialized meiotic membranes as nuclear segregation is being completed, thus triggering kinase activity at a specific place and time during this developmental program.
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