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Granule cells and Purkinje cells are the major populations of neurons in the cerebellum. Their specification depends on a combination of regional identity and spatiotemporal cues. These are conferred by patterning systems in the early embryo that determine anteroposterior and dorsoventral positional coordinates and an age-dependent signal (or signals) whose nature is obscure. While a number of important questions remain about the nature of cerebellar progenitor pools and their precise boundaries, a variety of fate-mapping and genetic T. Butts

What makes us human? That we walk upright, on two legs? That we have complex language? That we use tools? Or is it our ability to ask this question? For centuries, philosophers and scientists have all grappled with this fundamental question.

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