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LactobaciUus delbrueckii, strain 730 (ATCC 9649), is representative of a large group of lactic acid bacteria that is more fastidious in its nutritional requirements than are those organisms (e.g., LactobaciUs arabinosus, Strepcocus faecalis) commonly used for assay of vitamins and amino acids. Previous nutritional investigations with this organism (McNutt and Snell, 1950; Kitay et al., 1950) showed it to require pyridoxamine phosphate, thymidine, and an unsaturated fatty acid (e.g., oleic acid) for growth in a medium supplied with the usual vitamins and amino acids. The requirement forl thymidine was specific; vitamin B12, reducing agents, or

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