The clam Cyclina sinensis is one of the most important economic bivalves in China and the clam industry has developed rapidly in recent years (Liu et al., 2002;Wei et al., 2020). C. sinensis is readily obtained in coastal intertidal zone of East Asia, particularly in coastal muddy tidal and estuaries. The clam possesses lots of advantages as one of the main aquaculture species of shellfish, including rapid growth,
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