“…In light-exposed hot springs, microbial mats are dominated by phototrophic bacteria at temperatures between 42–74°C (70), and the microbial community composition and distribution of these mats exhibit niche differentiation along steep vertical gradients of light, water temperatures, and O 2 concentrations (26, 39, 45, 46). Previous studies have mostly focused on the importance of phototrophic organisms for primary production in the mats (10, 35, 57), and phototrophic bacteria have been shown to play important biogeochemical roles in facilitating oxygen production, sulfide consumption, and carbon fixation, among others (19, 21, 30, 36, 54, 56). Numerous chemotrophic organisms ( e.g.…”