“…A number of studies have examined biomass accumulation (Ewel et al, 1983;Chai, 1997;Hashimoto et al, 2000;Ohtsuka, 2001;Lawrence, 2005;Jepsen, 2006;Kenzo et al, 2010;Tanaka et al, 2021) and/or species diversity and composition (Ewel et al, 1983;Prajadinata, 1996;Lawrence et al, 2005;Wasli et al, 2011;Labrière et al, 2015;Karyati et al, 2018;Ulfah & Sulistyawati, 2018;Tanaka et al, 2021) in secondary forests of Borneo but they have largely focused on younger secondary forests (<20 years) with only a few studies on older forests (Okimori & Matius, 2000;Brearley et al, 2004;Wasli et al, 2011). When determining forest biomass, allometric equations are required as it is rarely possible, nor desirable, to directly measure and enumerate all the trees present through direct harvesting.…”