A consideration of the morphological aspects of the earliest modern humans in Europe (more than Ϸ33,000 B.P.) and the subsequent Gravettian human remains indicates that they possess an anatomical pattern congruent with the autapomorphic (derived) morphology of the earliest (Middle Paleolithic) African modern humans. However, they exhibit a variable suite of features that are either distinctive Neandertal traits and/or plesiomorphic (ancestral) aspects that had been lost among the African Middle Paleolithic modern humans. These features include aspects of neurocranial shape, basicranial external morphology, mandibular ramal and symphyseal form, dental morphology and size, and anteroposterior dental proportions, as well as aspects of the clavicles, scapulae, metacarpals, and appendicular proportions. The ubiquitous and variable presence of these morphological features in the European earlier modern human samples can only be parsimoniously explained as a product of modest levels of assimilation of Neandertals into early modern human populations as the latter dispersed across Europe. This interpretation is in agreement with current analyses of recent and past human molecular data.crania ͉ dentition ͉ Late Pleistocene ͉ postcrania T he evolutionary fate of the Neandertals has preoccupied paleoanthropologists with varying degrees of intensity for approximately a century, emerging with the human paleontological discoveries of the early 20th century and the resultant syntheses. Although present throughout the 20th century, considerations of the fate of the Neandertals have intensified in the past quarter-century starting with the paleontological emergence of Out-of-Africa models of modern human origins (1, 2) that combined the early appearance of modern human anatomy in Africa (2, 3) with the presence of equatorial anatomical patterns among the earliest modern Europeans (1). These interpretations were joined half a decade later by inferences from extant human molecular analyses, which rapidly polarized the field into two opposing models, best termed the Replacement (Out-of-Africa with total replacement of all other indigenous human populations) and Regional Continuity (variable regional transitions to modern human morphology among genetically interconnected human populations). † After two decades of debate, it is now recognized that these polarized models, Replacement (sensu stricto) vs. Regional Continuity (sensu stricto) are intellectually dead. Repeated attempts to refute one or the other model have shown that it is easy, on a global basis, to refute both scenarios with paleontological and/or extant human biological (both anatomical and molecular) data. It is time to move on from these models, and, particularly in the past half-decade, the emergent consensus model of modern human emergence is one of Out-of-Africa with temporally and geographically varying degrees of absorption of regional late archaic populations [the Assimilation Model (10)]. The issue is no longer whether human populations outside of the African ...