“…Epistatic interactions between genetic variation in the nuclear and cytoplasmic genomes, cytonuclear interactions, have been found in most species where they have been empirically tested, including plants, insects, fungi, and animals (Burke et al, 1998;Roubertoux et al, 2003;Tao et al, 2004;Zeyl et al, 2005;Dowling et al, 2007Dowling et al, , 2010Etterson et al, 2007; The proportion of genetic variance explainable by various genetic factors in this population, as measured using type III sums of squares, for aliphatic glucosinolates (red online, left), indole glucosinolates (blue online, center), and growth (green online, right). Error bars show SE, and there are 15 aliphatic glucosinolate traits, six indolic glucosinolate traits, and seven growth traits.…”