“…Previous studies have revealed that plant C, N and P contents relate to numerous abiotic and biotic factors, including plant growth stage (Elser et al 2000), grazing (Zheng et al 2012, Zhang et al 2014, and plant functional traits (Reich et al 1999). As such, studies often investigate the changes in N, C and P in accordance with a single ecological factor at the regional scale or in one habitat (Niu et al 2013.…”