Eine neue Unterart vonPhacops (Phacops) sparsinodosus Struve 1970 aus dem Mittel-Devon von Marokko

Abstract

Eine neue Unterart von Phacops (Phacops) sparsinodosus STRUVE 1970 aus dem Mittel-Devon von Marokko Mit 8 Abbildungen GUNNAR SCHRAUT Kurzfassung Aus dem tiefen Mittel-Devon (tiefes Eifelium) des Profils Zguid III ira sª Antiatlas (Marokko) wird die neue Unterart Phacops (Phacopa9 sparsinodosus struvei n. ssp. aufgestellt und stratigraphisch mit Hilfe ron Ammonoideen und Trilobiten eingehfingt. Der Fund eines vollst~indigen Schalenexemplars von Phacops (Phacops) sparsinodosus gallicus (STRuVE 1982) aus den Grauwacken-Schiefern der Assise de Bure, Couvinien Col, Grenz-Bereich Unter-Devon/Mittel-Devon, erfordert eine Neudarstellung dieses Taxons und erm6glicht einen guten Vergleich mit den bisher bekannten Unterarten. Durch das Vorkommen von Fidelites sp. und Thysanopeltis speciosa in Marokko und von Pinacitesjugleri in Belgien ist die stratigraphische Einstufung beider Unterarten in das untere Mittel-Devon (tiefes Eifelium) gesichert. Abstract [A new subspecies of Phacops (Phacops) sparsinodosus STRUVE 1970 from the Middle Devonian of Morocco.] The new subspecies Phacops (Phacops) sparsinodosus struvei n. ssp. is based on material from the Zguid fil section (southern part of the Antiatlas, Morocco). Age correlations were made using ammonoids and trilobites. A nearly complete specimen of Phacops (Phacops) sparsinodosus gallicus (STRuVE 1982) with exce[lent shell preservation is described. Ir was found in the ,,Grauwacken-Schiefern" of the Assise de Bure in Belgium (Couvinien Col), type-locality of the taxon.It is now possible to compare all subspecies of Ph. (Ph.) sparsinodosus with the same level of preservation. The trilobite records together with the occurrences ofFidelites sp., Thysanopeltis speciosa (Morocco) and Pinacites jugleri (Belgium) indicate ah approximate stratigraphic position within the lower part of the Eifelian (Middle Devonian).

The phacopine trilobite genera Morocops Basse, 2006 and Adrisiops gen. nov. from the Devonian of Morocco

“…Regardless of the identities of the various paratypes, the holotype of M. struvei bears all of the diagnostic features of Barrandeops as according to McKellar & Chatterton's (2009) and is, above all, similar to stratigraphically young species of the group (see below). Furthermore, we disagree with the previous viewpoint of Chatterton et al (2006) who considered all of the types figured by Schraut (2000b) Khaldi et al (2016). McKellar & Chatterton (2009) were conservative in only assigning to Barrandeops species described by themselves and by Chatterton et al (2006) from Morocco.…”

“…8m-o) is in more than those ways dissimilar from the other types and rejected Schraut's claim that the differences observed should all be attributed to ontogeny. McKellar & Chatterton (2009) inadvertently took Basse (2006) to have corroborated Schraut's (2000b) ontogenetic suggestion and advanced arguments as to why the differences observed were unlikely to occur within a single species (i.e., in fact, they agreed fully with Basse 2006). McKellar & Chatterton (2009, p. 21) stated that they coded M. struvei for cladistic analysis based on, "the large form (the type specimen and paratypes sharing its thick sclera)", although they did not specify these paratypes nor did they provide arguments for their decision.…”

“…These contrasting opinions arose because the type species of Morocops, M. struvei, has been subject to different interpretations. Schraut (2000b) described M. struvei as a subspecies of Geesops sparsinodosus (Struve, 1970) while noting what he considered to be minor ontogenetic variations among his type series. These are changes in the divergence of the glabellar axial furrows, the lateral inclination of the eyes and the size and distribution of cephalic sculpture.…”

“…Schraut (2000b, fig. 8d-f) recorded an exceptionally well-preserved, enrolled specimen of that subspecies that differs merely from the types of M. lebesus figured by Chatterton et al (2006, pl.…”

Morphological trends and new species of Cyphaspis (Trilobita, Otarioninae) in the Devonian of Morocco, Turkey, Germany and Belgium

“…1, point 2). Trilobites (data for Morocco here and further below are based on field notes by DH and the late H. Prescher) from ZGEE1, ZGEE2 (upper Emsian): Acastoides zguilmensis Chatterton et al, 2006, Austerops menchikoffi (Le Maître, 1952 Chatterton et al, 2006, following Khaldi et al (2016], Cyphaspis eberhardiei, Diademaproetus praecursor Alberti, 1969, Erbenochile erbeni (Alberti, 1981), Gandlops tamnrherta (Chatterton et al, 2006), Gerastos marocensis Chatterton et al, 2006 [herein regarded as a distinct species instead of a subspecies for reasons discussed by van Viersen & Prescher (2010) with regard to other Gerastos species], Hollardops mesocristatus (Le Maître, 1952), Koneprusia dahmani Chatterton et al, 2006, Leonaspis haddanei Chatterton et al, 2006, Leonaspis spinicurva Chatterton et al, 2006, Morocops granulops (Chatterton et al, 2006) [the type series of Phacops (Phacops) sparsinodosus struvei Schraut, 2000 (type species of Morocops Basse, 2006) from the Timrhanrhart Formation at Foum Zguid encompass at least two distinct phacopid species; the holotype (Schraut, 2000, fig. 8g-i Morzadec, 2001, Wenndorfia calva (Chatterton et al, 2006), undescribed genera and species of Scutelluinae.…”