“…Ciliated cells underlie several annelid sensory modalities (including vision, mechanoreception, and chemoreception) that are processed by the central nervous system (Mill, 1978). Among these senses, chemoreception modulates a variety of ecologically important behaviors including spawning (Hardege et al, 1996;Hardege and Bentley, 1997;Hardege, 1999), possibly spermataphore transfer in spionids (Rice, 1978(Rice, , 1991, larval settlement (reviewed by Qian, 1999), juvenile postsettlement movement (Woodin et al, 1995), and feeding (Copeland and Weiman, 1924;Rullier, 1950;Ferner and Jumars, 1999;Kihslinger and Woodin, 2000;Riordan and Lindsay, 2002;Mahon and Dauer, 2005).…”