“…In addition to growth traits, wood traits, such as wood density and strength properties, have been considered essential for sustainable wood utilization. In tree breeding for wood quality, non-destructive and semi-destructive testing methods are applied to evaluate wood traits to decrease the cost and increase efficiency (Chen et al, 2010;Wu et al, 2011;Hidayati et al, 2013Hidayati et al, , 2019Hai et al, 2015). For example, nondestructive testing, stress-wave velocity (Wu et al, 2011;Hidayati et al, 2013Hidayati et al, , 2019Hai et al, 2015), and semi-destructive testing, Pilodyn penetration (Chen et al, 2010;Wu et al, 2011;Hidayati et al, 2013Hidayati et al, , 2019Hai et al, 2015) can evaluate Young's modulus and wood density of stems in standing trees, respectively.…”