Land snails are ideal materials for paleoclimate studies (Goodfriend, 1992;Wang et al., 2016;Wu et al., 2018); this is because they have the advantages of being widely distributed, abundant, and well preserved in sedimentary succession strata. Additionally, they are relatively sensitive to climate changes. So far, research on land snails has included inferring the environmental conditions under which land snails survive by identifying faunal assemblages and living habits of each species (Gittenberger & Goodfriend, 1993;Wu & Li, 2008) and reconstructing the paleoclimates by analyzing stable isotopes of land snail shells (Colonese