“…Consequently, the driving principles of phenotypic variation in individual-environment interactions are in the focus of current research (e.g., Chevin, Lande, and Mace 2010;Ameline et al 2018;Hein et al 2018). However, knowledge about specific spatial and temporal variations of life history traits remains scarce in invertebrates, especially in cold environments (Homburg et al 2013;Høye and Sikes 2013;Ameline et al 2017). Body size is one of the key features of life history traits research in ectotherms (Angiletta et al 2004;Chown and Gaston 2010;Høye and Hammel 2010), because it is considered a proxy for fecundity, body condition, and survival (e.g., Hodkinson 2005;Bowden, Høye, and Buddle 2013;Penell, Raub, and Höfer 2018).…”