“…This approach was widely employed for direct, quantitative paleo-temperature reconstructions including examples from the Precambrian-Cambrian (Meng et al, 2011b), Silurian (Losey and Benison, 2000), Permian (Benison and Goldstein, 1999; Zambito and Benison, 2013), Paleogene (Meng et al, 2013;Zhao et al, 2014), Quaternary (Lowenstein et al, 1998(Lowenstein et al, , 1999 and modern settings (Roberts and Spencer, 1995).…”