A Late Eocene foraminiferal fauna of free specimens is recorded from Cascade Seamount, a feature of volcanic origin, whose age and fauna were previously studied only in thin-section. The fauna is of Late Eocene (P15) age because of the presence of Globigerapsis index, Globigerapsis rubriformis, Subbotina linaperta, Subbotina angiporoides and Chiloguembelina cubensis. Many typical Eocene indicators, such as Hantkenina, Pseudohastigerina and key globorotaliids, are absent. The age is a little older than that estimated for previous samples from the seamount. The fauna is dominated (78%) by benthic species, especially species of Cibicides that have greater affinities with New Zealand faunas than with coeval southern Australian faunas. The sediment and its fauna accumulated near the lower limits of wave activity, off the coast of a volcano that probably stood at least 400 m above sea-level. The location has subsided nearly 1000 m over the past 40 million years to its present water depth of ~1000 m, suggesting that the entire East Tasman Plateau was approximately 1000 m above its current depth at the time. Conditions were cool temperate and the fauna lacks any warm-water indices, consistent with a palaeolatitude of 55-60°S, but also with cool waters at a time of oscillating conditions in southern Australia and also in Antarctica. There is essentially no infauna and the waters were highly oxygenated in the turbulent zone.