“…It is widely known that climatic fluctuations during the Pleistocene strongly impacted the distribution and genetic structure of plant and animal species in both Northern and Southern Hemispheres (Hewitt, 2004;Lu, Heckel, Liang, & Zhang, 2013;Song et al, 2009). Although China experienced less severe glaciations during this period relative to Europe and North America (López-Pujol, Zhang, Sun, Ying, & Ge, 2011;Qian & Ricklefs, 1999Rost, 2000), climatic oscillations nonetheless affected the genetic diversity and evolution of many extant species in the region (Feng, Zheng, & Gong, 2016;Song et al, 2009;Zhou et al, 2010). Recent phylogeographic surveys in China indicate that during Pleistocene glaciations montane species might frequently have persisted in topographically complex regions that served as refugia, rather than being restricted only to lower latitudes Liu et al, 2013;Qiu, Fu, & Comes, 2011;Song et al, 2009;Zhou et al, 2010).…”