“…In the past, determining the abundance of LOC in the seabed has been difficult due to limited sampling resolution and short (on the order of a few months) LOC residence times (Hammond et al, 1996). No direct measurements of LOC have been developed; only proxies for the "reactive" organic carbon content such as Total Hydrolyzable Amino Acids (THAA; e.g., Grémare et al, 2003;Garcia et al, 2010;Sañé et al, 2013b), biopolymeric organic carbon (e.g., Fabiano et al, 1995;Fabiano and Pusceddu, 1998), porewater diagenetic reactions (Arndt et al, 2013;Komada et al, 2016), or chlorophyll-a (e.g., Stephens et al, 1997;Sañé et al, 2011b). Common problems with these approaches include representation of only a small portion of the total LOC pool (such as in the case of chlorophyll-a or THAA) or the approaches also include older, less reactive organic carbon that has limited nutritional value to the benthos (e.g., biopolymeric organic carbon).…”