“…Therefore, medium sized snails are best suited to withstand desiccation on the soil surface because of a more favourable aperture size/weight ratio (Hira, 1968;Diaw et al, 1988). Secondly, humid soils offer better survival chances (Ohlweiler & Kawano, 2001;Kalinda et al, 2018a) although aerobic conditions remain a prerequisite (Chu et al, 1967a;Coles, 1969;Betterton, 1984;Ebele & Smith, 1990;Thomas & Daldorph, 1991). This explains why survival rates are lower in wet mud and places exposed to sunlight (Chu et al, 1967c;Hira, 1968;Van Aardt & Steytler, 2007) and why they increase when IH snails bury in moist mud (Chu et al, 1967a;Kalinda et al, 2018b).…”