Abstract:In spite of recent interest, the various species belonging to the genera Heminautilus SPATH, 1927, and Josanautilus MARTÍNEZ &GRAUGES, 2006, have never been the object of a general revision. More than 160 specimens belonging to various species of these two genera were studied, with the aim of identifying the specific characteristics and the stratigraphical and paleogeographical distribution of the various species which compose them, as well as to propose a phylogeny for the group. This study confirms the non-synonymy of H. saxbii (MORRIS, 1848) and H. lallierianus (ORBIGNY, 1841), establishes that H. tejeriensis MARTÍNEZ &GRAUGES, 2006, and H. verneuilli (VILANOVA, 1870) represent junior synonyms of H. saxbii (MORRIS) and suggests the creation of the species H. ? japonicus sp. nov. The origin of the genus Heminautilus SPATH from Pseudocenoceras SPATH, 1927, proposed by TINTANT and the origin of Josanautilus MARTÍNEZ & GRAUGES from Heminautilus SPATH are both confirmed.Key Words: Nautiloidea; Cenoceratidae; Heminautilus; Josanautilus; Early Cretaceous; Barremian; Aptian.Citation: BAUDOUIN C., DELANOY G., MORENO-BEDMAR J.A., PICTET A., VERMEULEN J., CONTE G., GONNET R., BOSELLI P. & BOSELLI M. (2016).-Revision of the Early Cretaceous genera Heminautilus SPATH, 1927, and Josanautilus MARTÍNEZ & GRAUGES, 2006 (Nautilida, Cenoceratidae). Madrid, vol. 16, no. 5, Résumé : Révision des genres Heminautilus SPATH, 1927, et Josanautilus MARTÍNEZ & GRAU-GES, 2006 (Nautilida, Cenoceratidae), du Crétacé inférieur.-Malgré des travaux récents, les diffé-rentes espèces appartenant aux genres Heminautilus SPATH, 1927, et Josanautilus MARTÍNEZ & GRAUGES, 2006, n'ont jamais fait l'objet d'une révision d'ensemble. Plus de 160 spécimens appartenant à diffé-rentes espèces de ces deux genres ont été étudiés dans le but de cerner les caractères spécifiques et la répartition stratigraphique et paléogéographique des différentes espèces qui les composent, ainsi que de proposer un schéma phylogénétique de ce groupe. Cette étude a permis en particulier de confirmer la (1980, 1985, 1989, 2007, 2010), MARTÍNEZ & GRAUGES (2006) and DELANOY et al. (2012) The classification of SPATH SPATH (1927) is the first author to propose a classification of Nautiloidea, based purely on morphological criteria. Thus he classifies the genus Heminautilus SPATH, 1927, in the Family Paracenoceratidae SPATH, 1927, which gathers all the forms with a differentiated ventral region and a suture line containing a well developed ventral lobe. However, SPATH (1927, p. 25) himself recognizes the polyphyletic character of this family. The works of TINTANT (TINTANT & KABAM- BA, 1983;TINTANT, 1993) will later show that this classification does not actually take into account phyletic data and that numerous identical morphological characteristics appear in an iterative way in various lineages. The classification of SPATH was used, however, by DZIK (1984, p. 181) who presented a phylogenetic outline of the Family Paracenoceratidae SPATH.
The classification of KU...