In this paper, we present an infrared transparent frequency selective surface (ITFSS) based on iterative metallic meshes, which possesses the properties of high transmittance in infrared band and band-pass effect in millimeter wave band. Cross-slot units are designed on the iterative metallic meshes, which is composed of two same square metallic meshes with a misplaced overlap. In the infrared band of 3–5 μm, the ITFSS has an average transmittance of 80% with a MgF2 substrate. In the millimeter wave band, a transmittance of −0.74 dB at the resonance frequency of 39.4 GHz is obtained. Moreover, theoretical simulations of the ITFSS diffractive characteristics and transmittance response are also investigated in detail. This ITFSS may be an efficient way to achieve the metamaterial millimeter wave/infrared functional film.