A new structural study of a D
2
-M
2
tectono-thermal structure in SW Iberia (Ponte de Sor-Seda gneiss dome) characterized by a spatial distribution of telescoping isograds providing a record of Buchan-type metamorphic conditions is presented. It comprises an infrastructure made up of the Lower Gneiss (LGU) and the Intermediate Schist (ISU) units separated by early D
2
ductile extensional shear zones. The LGU and the ISU are composed of Ediacaran-Cambrian rocks that experienced the M
2
highest-grade metamorphic conditions (amphibolite facies). Late Ediacaran-Early Terreneuvian and Late Miaolingian-Early Furongian protolith ages for LGU (496 ± 3 Ma) and ISU (539 ± 2 Ma) orthogneisses are reported. The superstructure made of Cambrian-Devonian rocks (Upper Slate Unit. USU) deformed under M
2
greenschist facies, tectonically overlies the ISU across a D
2
extensional shear zone. Kinematic criteria associated with D
2
-M
2
fabrics indicate top-to-ESE-SE sense of shear. A late-D
2
brittle-ductile high-angle extensional shear zone (Seda shear zone) crosscut the gneiss dome. D
3
upright folds, thrusts and transpressive shear zones caused the steepening of D
2
structures and the local crenulation of S
2
foliation. The Mississippian D
2
-M
2
event recorded in the OMZ may be regarded as a regional scale phenomenon that markedly influenced the crustal architecture of North Gondwana during the assembly of Pangea.
Supplementary material at
https://doi.org/10.6084/m9.figshare.c.6828875