“…Low yields of upland rice are often attributed to sub-optimal crop management practices, limited availability of high-yielding varieties, and abiotic and biotic stresses (some of which might be associated with land use pressure) (Becker & Johnson, 2001;Diagne et al, 2013;Niang et al, 2017;Saito, Linquist, Keobualapha, et al, 2006;Saito et al, 2013;Tanaka et al, 2017). Typical major abiotic and biotic stresses include cold stress in highlands (Ahmadi et al, 2004;Raboin et al, 2014), drought (Kijima et al, 2011;Niang et al, 2017Niang et al, , 2018Saito et al, 2013Saito et al, , 2017, soil-related problems including acidity and N and P deficiency (Asai et al, 2009;Haefele et al, 2014;Okada & Fischer, 2001;Saito, Linquist, Keobualapha, et al, 2006;Saito et al, 2007Saito et al, , 2013, weeds (Becker & Johnson, 2001;, and soil-borne pests such as nematodes (Balasubramanian et al, 2007).…”