“…Many studies have considered all seed properties to be offspring traits (e.g., Byers et al, 1997), and accordingly have interpreted the seeds of each mother plant as half-sibs (e.g., Carles et al, 2009). In most of these cases, including almost all the studies reporting heritability for seed mass in conifer trees (Matziris, 1998;Bilir et al, 2008;Sivacioglu and Ayan, 2008;Carles et al, 2009), the reported heritability estimates were, in fact, clonal repeatability estimates. The statistical analyses used have not accounted, however, for the fact that seeds of a given mother plant are, indeed, non-independent repeated measures within the same subject (Carles et al, 2009).…”