Abstract-This paper reports the theory and implementation of a decentralized navigation system that enables simultaneous single-beacon navigation of multiple underwater vehicles. In single-beacon navigation, each vehicle uses ranges from a single, moving reference beacon in addition to its own inertial navigation sensors to perform absolute localization and navigation. In this implementation the vehicles perform simultaneous communication and navigation using underwater acoustic modems, encoding and decoding data within the acoustic broadcast. Vehicles calculate range from the time of flight of asynchronous acoustic broadcasts from the reference beacon. Synchronous clocks on the reference beacon and the vehicles enable the measurement of one-way travel-times, whereby the time of launch of the acoustic signal at the reference beacon is encoded in the acoustic broadcast and the time of arrival of the broadcast is measured by each vehicle. The decentralized navigation algorithm, running independently on each vehicle, is implemented using the information form of the extended Kalman filter and has been previously shown to yield results that are identical to a centralized Kalman filter at the instant of each range measurement. We summarize herein the architecture and design of the acoustic communications (Acomms) system consisting of an underwater acoustic modem, synchronous clock, and the software necessary to run them, and salient results from the validation of the decentralized information filter using a simulated data set.