Ab stractThe oc cur rence of Pteridium spores -the com mon fire-adapted plant -was ob served in Po land's Pleis to cene (three interglacials and nu mer ous interstadials) and Ho lo cene pol len se quences. Un til the on set of the mid dle Ho lo cene, bracken was re corded rel a tively rarely. This marks cli ma tic con di tions with spo radic wildfires, fol lowed by quick removal of the clones (or main te nance of the clones in a sup pressed state) in the sub se quent, post fire successional stages. In each inter gla cial we can also iden tify short pe ri ods (most of ten syn chro nous) of some what higher fre quency of Pteridium, in di cat ing a pos si ble in crease in nat u ral fires as an ef fect of stormy con di tions. These short pe ri ods are placed mainly within phases dom i nated by co nif er ous wood lands. The very high con tent of bracken found from a reanalysis of the long Saalian se quence at Ossówka as three reg u lar cul mi na tions just above three non-tree phases, east ern Po land is the unique ex cep tion. To ex plain this we found the close mod ern an a logue of the above phe nom e non in the Alas kan re gion, at the bound ary be tween the tun dra and the bo real zone where in a very nar row for est zone, espe cially with the con ti nen tal sig na ture, light ning-ini ti ated fires are very fre quent (the edge ef fect). Pleis to cene re cords of bracken dur ing for est pe ri ods might in di cate that thun der storms and light ning strikes were re spon si ble for its higher con tent. This is not to ex clude the pos si bil ity that inter gla cial fires were set by Palaeo lithic hu mans. How ever, it is more likely that the wildfires were uti lized and to some ex tent con trolled, es pe cially at Ossówka, where the palaeolake ex isted for a long time af ter the inter gla cial; and this surely at tracted the at ten tion of game and hu mans. Our in ves ti gations show that rare, more abun dant Pteridium in the Pleis to cene se quences can be traced through out the cor re sponding pe ri ods even at dis tant sites. This might be in di rect ev i dence of a cli ma tic pat tern that pro mote stormy con di tions and fires at that time. In the Ho lo cene, dif fer ent fac tors seem to be re spon si ble for the long-term dy nam ics in Pteridium clones. Both in Po land and in ad ja cent ar eas bracken peaked in the mid dle Ho lo cene from 8000 to 5000 BP, when this ter ri tory was oc cu pied by de cid u ous wood lands. There is agree ment that this is due to burn ing of for ests by the hunter-gath erer so ci et ies of the Late Mesolithic. How ever, in spite of in creased clear ings in the sub se quent phases bracken sub stan tially de creased in abun dance. This means first of all that fires rather than clear ings were re spon si ble for the rise in Pteridium spores in the Ho lo cene pol len se quences.Key words: bracken spores, fires, Qua ter nary, Po land
IN TRO DUC TIONPteridium aquilinum is a com mon plant grow ing on every con ti nent ex cept in Arc tic/Ant arc tic re gions. For many rea sons -first of a...