The topology of natural fracture networks is inherently linked to the structure of the fluid velocity field and transport therein. Here we study the impact of network density on flow and transport behaviors. We stochastically generate fracture networks of varying density and simulate flow and transport with a discrete fracture network model, which fully resolves network topology at the fracture scale. We study conservative solute trajectories with Lagrangian particle tracking and find that as fracture density decreases, solute channelization to large local fractures increases, thereby reducing plume spreading. Furthermore, in sparse networks mean particle travel distance increases and local network features, such as velocity zones where flow is counter to the primary pressure gradient, become increasingly important for transport. As the network density increases, network statistics homogenize and such local features have a reduced impact. We quantify local topological influence on transport behavior with an effective tortuosity parameter, which measures the ratio of total advective distance to linear distance at the fracture scale; large tortuosity values are correlated to slow-velocity regions. These large tortuosity, slow-velocity regions delay downstream transport and enhance tailing on particle breakthrough curves. Finally, we predict transport with an upscaled, Bernoulli spatial Markov random walk model and parameterize local topological influences with a novel tortuosity parameter. Bernoulli model predictions improve when sampling from a tortuosity distribution, as opposed to a fixed value as has previously been done, suggesting that local network topological features must be carefully considered in upscaled modeling efforts of fracture network systems. and topology, and the corresponding flow field. The flow field within an individual fracture is typically highly correlated, commonly causing solute velocity to display persistent, low variability behavior over the in-fracture scale; consequently, the greatest Lagrangian accelerations occur at fracture intersections . As the fracture density increases, solute encounters more intersections on average, and the velocity correlation scale decreases. Furthermore, strong preferential flow paths form within interconnected networks of large fractures and channel a significant portion of mass, enabling solute to persist at high velocities for distances greater than the single fracture scale Sherman et al., 2019). This channelization becomes enhanced in sparse networks, where particles encounter fewer intersections, enabling them to persist on single fractures for longer distances. Resolving all these intranetwork features in 3-D DFN models is still computationally costly, and so upscaled modeling approaches, which account for network variability through effective parameter schemes, while maintaining a parsimonious framework, present an attractive alternative. However, how to properly parameterize network properties, such as velocity correlation and geometry, and inco...