Abstract. Current theories of F-layer storms are discussed using numerical simulations with the Upper Atmosphere Model, a global self-consistent, time dependent numerical model of the thermosphere±iono-sphere±plasmasphere±magnetosphere system including electrodynamical coupling e ects. A case study of a moderate geomagnetic storm at low solar activity during the northern winter solstice exempli®es the complex storm phenomena. The study focuses on positive ionospheric storm e ects in relation to thermospheric disturbances in general and thermospheric composition changes in particular. It investigates the dynamical e ects of both neutral meridional winds and electric ®elds caused by the disturbance dynamo e ect. The penetration of short-time electric ®elds of magnetospheric origin during storm intensi®cation phases is shown for the ®rst time in this model study. Comparisons of the calculated thermospheric composition changes with satellite observations of AE-C and ESRO-4 during storm time show a good agreement. The empirical MSISE90 model, however, is less consistent with the simulations. It does not show the equatorward propagation of the disturbances and predicts that they have a gentler latitudinal gradient. Both theoretical and experimental data reveal that although the ratio of [O]/[N 2 ] at high latitudes decreases significantly during the magnetic storm compared with the quiet time level, at mid to low latitudes it does not increase (at ®xed altitudes) above the quiet reference level. Meanwhile, the ionospheric storm is positive there. We conclude that the positive phase of the ionospheric storm is mainly due to uplifting of ionospheric F 2 -region plasma at mid latitudes and its equatorward movement at low latitudes along geomagnetic ®eld lines caused by large-scale neutral wind circulation and the passage of travelling atmospheric disturbances (TADs). The calculated zonal electric ®eld disturbances also help to create the positive ionospheric disturbances both at middle and low latitudes. Minor contributions arise from the general density enhancement of all constituents during geomagnetic storms, which favours ion production processes above ion losses at ®xed height under daylight conditions.