“…Natural habitats often increase the diversity and abundance of natural enemies (Bianchi et al, 2006;Chaplin-Kramer, O'Rourke, Blitzer, & Kremen, 2011). Area (Fahrig, 2003), isolation (Kruess & Tscharntke, 2000), permeability (i.e., perimeter/area ratio, (Stoner & Joern, 2004;Wu, 2007)), and proximity to noncrop habitats (Clough, Kruess, Kleijn, & Tscharntke, 2005;González, Salvo, & Valladares, 2015;Tscharntke, Gathmann, & Steffan-Dewenter, 1998) influence arthropod densities and distribution, and their beneficial effects on crops. Temporal dynamics is also important, as movements of natural enemies between natural fragments and crops change in direction and intensity (Macfadyen et al, 2015;Rand, Tylianakis, & Tscharntke, 2006).…”