Petrological and δ 13 C analyses were under taken on contiguous specimens of coal and intercalated minor organicrich clastic sedi ments collected from coal seams spanning the CretaceousPaleogene boundary in the Alberta and Saskatchewan portions of the Western Interior Basin. The generally high smectite content of the coal suggests that the original mires were largely small, discon nected, and rheotrophic, readily receiving abundant waterborne detrital clastic mate rial of largely volcanic origin. Nevertheless, using the distinctive claystone that marks the CretaceousPaleogene boundary as a regional datum , it is possible to correlate cycles in the vitrinite and inertinite compo sition of the coals over >500 km. Estimates of peat accumulation rates suggest that the cycles in vitrinite and inertinite composi tion represent regional, cyclic fluctuations in wildfire and oxi da tion of the peatlands and overlying canopy at a frequency of hundreds to thousands of years. The likely causes of these fluctuations were cyclic, regionalscale changes in temperature. The Cretaceous Paleogene boundary event occurred early during a phase of gradually increasing tem perature and/or decreasing rainfall, but peak wildfire and desiccation of peat occurred up to 14,000 yr later than the Cretaceous Paleogene boundary, and the mires did not experience significant water stress in the im mediate aftermath of the extinction event. A persistent, 1.5‰-3.0‰ negative δ 13 C excur sion occurs across the CretaceousPaleogene boundary, but it cannot be readily separated from four, further negative excursions later in the earliest Danian. The negative carbon isotope excursion linked to the Cretaceous Paleogene boundary began a few hundred years before the event itself, and recovery occurred within 21 k.y., and possibly in as little as just a few thousand years, consistent with recently calibrated shallowmarine δ 13 C records. Hence, the atmospheric and surface ocean carbon pools were coupled at this time. The absence of evidence for catastrophic change in the climatic regime at the time of the CretaceousPaleogene extinction in these mires supports the notion that the negative shift in atmospheric δ 13 C was brought about by changes in the δ 13 C composition of the sur face ocean. This is consistent with the greater magnitude of extinction experienced by ma rine fauna relative to the terrestrial realm.