“…Key sequence stratigraphic surfaces (e.g., sequence boundaries, transgressive surfaces) are frequently marked by substrate-controlled ichnofossil assemblages (e.g., Glossifungites Ichnofacies; Savrda, 1991a;MacEachern et al, 1992;Pemberton et al, 2000;Buatois and Encinas, 2006;Rodríguez-Tovar et al, 2006;Abdel-Fattah et al, 2016), and vertical changes in ichnofossil assemblages can be used to recognize the signal of sea-level change (e.g., shallowing-upward patterns manifested in parasequences; Savrda, 1991bSavrda, , 1995Brett, 1998;Fielding et al, 2006;Rodríguez-Tovar et al, 2010;Paranjape et al, 2014). Less commonly appreciated among sedimentary geologists, except perhaps those working mainly in the Paleozoic, is the potential use of trace fossils in ichnostratigraphy, a variant of biostratigraphy that aids in determining stratigraphic age of sedimentary sequences (Crimes, 1969(Crimes, , 1970(Crimes, , 1975Seilacher, 1970Seilacher, , 1992Seilacher, , 1994Seilacher, , 1996Seilacher, , 2000Seilacher, , 2007Rindsberg and Martin, 2003;Baldwin and Strother, 2004;Mángano et al, 2001;Mángano and Droser, 2004;Mángano et al, 2005;Brandt et al, 2010).…”