Prehistoric societies from around the world constructed monumental mounded architecture (earthen pyramids) for a variety of functions, including the foundation for temples or a leader's residence, community stages and cemeteries. Flat-topped earthen mounds often have complicated histories where the function, size, orientation and summit architecture varied throughout time. This paper presents the results from a ground-penetrating radar (GPR) survey conducted on the summit of Mound A, the largest prehistoric Native American mound at Ocmulgee National Monument located in central Georgia, USA. Our study indicates that performing depth-slice analyses of flat-topped mounds can effectively map successive construction stages over distinct periods of archaeological prehistory. The GPR data show distinctive low-amplitude, discontinuous stratigraphic variations that we interpret to be related to mound fill, which are interrupted by high-amplitude coherent summit reflections. We also maintain sufficient vertical and horizontal resolution to identify summit architecture on earlier mound-use episodes, which is imaged by distinct reflection geometry as patterned linear, square and circular high-amplitude events in GPR depth-slices. The authors recorded four possible mound summits, the western expansion of the mound, and the shifting location and shape of summit architecture; in addition to resolving a discrepancy regarding the location of early excavation units from the 1930s. Shallow geophysical data indicate a decline in the volume of material used over the course of mound construction, and by inference, a decline in the size of the labour force used to construct each stage. We conclude that the power and influence of Ocmulgee's early leadership subsided over the course of Mound A's use, and may have been contested by an emerging faction.