“…Within the Suborder Typotheria , the family Hegetotheriidae (late Oligocene–Early Pleistocene) includes placental mammals of small to medium size, with hypertrophied, hypselodont, and procumbent first incisors, hypselodont cheek teeth, lower molariforms with a deep labial groove, P2–M3 imbricated, and loss of ectoloph plications on upper molars (Cerdeño & Reguero, 2015; Kramarz & Paz, 2013; Seoane & Cerdeño, 2019). Hegetotheriidae currently comprises 11 genera distributed in two subfamilies: Hegetotheriinae , recognized as paraphyletic in most recent phylogenetic analyses (except for Billet, 2011; Seoane et al, 2017, and this work), and Pachyrukhinae , a well-established monophyletic group (Billet, 2011; Cerdeño & Reguero, 2015; Croft & Anaya, 2006; Kramarz & Bond, 2017; Reguero & Prevosti, 2010; Seoane & Cerdeño, 2019; Solórzano et al, 2023; Vera, 2019).…”