“…The overall distribution of the five basal genera (Margaritaria, Plagiocladus, Lingelsheimia, Heterosavia and Flueggea) is highly relictual. For example, Flueggea has a center of its range in the Old World tropics, but also has fragmentary distributions in the Iberian Peninsula, Turkey, West Indies, Ecuador and north-eastern Brazil (Webster 1984;Gemici 1993), which are most likely regarded as relicts (there is a single apparent dispersal to Hawaii; Hayden 1987). Similarly, Plagiocladus, Lingelsheimia and Heterosavia have highly restricted distributions in West Africa, Africa and Madagascar, and the West Indies, respectively (Table 1; Govaerts et al 2000;Radcliffe-Smith 2001;Lebrun & Stork 2006;Hoffmann 2008), suggestive of range contractions from once broader distributions.…”