“…The sister group to Asilidae is still debated in the Diptera literature and six alternative hypotheses are: (1) Asilidae and Apioceridae are sister taxa (Hull, 1962;Yeates, 1994); (2) either Apioceridae or Mydidae is the sister group to Asilidae (Hennig, 1973); (3) Asilidae and Mydidae form a sister taxon (Wood, 1981); (4) Asilidae is the sister taxon to either Apioceridae and Mydidae or Scenopinidae and Therevidae (Woodley, 1989); (5) Apioceridae and Mydidae form the sister group to Asilidae (Yeates and Irwin, 1996); and (6) Asilidae is the sister taxon to the remaining Asiloidea except Bombyliidae (Yeates, 2002;Bybee et al, 2004). Hermann (1909), in describing the first southern African Apioceridae in the genus Ripidosyrma Hermann, 1909, discussed the relationships of Apioceridae, Asilidae, and Mydidae and concluded that the Asilidae is most closely related to Apiocera Westwood, 1835, i.e., Australian Apioceridae, based on certain characters of the wing venation.…”