The Nanushuk Formation (Albian–Cenomanian) crops out over much of the central and western North Slope of Alaska, varying from ≈1500 to ≈250 m thick from west to northeast. The Nanushuk Formation records an inter-tonguing succession of marine and nonmarine conglomerate, sandstone, mudstone, and coal. These rock units comprise the Kukpowruk and Corwin formations of the former Nanushuk Group, respectively. Work presented here is centered in the foothills of the DeLong Mountains along the Kukpowruk River, from an area west of Igloo Mountain in the Coke Basin to the Barabara Syncline, approximately 80 km to the north. A radiometric date recovered from a tuff in our study area suggests a Cenomanian age for at least some of these rocks. Outcrops along the Kukpowruk River contain a well-preserved fossil flora previously recovered from marine, marginal marine, and terrestrial sediments. Our own work focuses on detailed measured sections of terrestrial rocks, interpretation of sedimentary facies and facies associations, and documentation of fossil vertebrates. Eight facies associations are identified in the study area that together are interpreted to represent meandering fluvial and upper delta plain environments. Plant fossils are common and include standing tree trunks up to 58 cm in diameter at some locations. Approximately 75 newly discovered tracksites, and a heretofore unknown, rich fossil vertebrate ichnofauna, are present. The ichnofaunal assemblage includes evidence of small and large theropod dinosaurs (including birds) and bipedal and quadrupedal ornithischian dinosaurs. Approximately 15% of the dinosaur ichnofauna record is represented by fossil bird tracks. Wood fragments from the Nanushuk Formation were analyzed for their carbon isotopic composition to relate δ13C to mean annual precipitation. Samples averaged −26.4‰ VPDB, suggesting an average MAP of 1412 mm/year. This record of increased precipitation in the Nanushuk Fm. during the mid-Cretaceous provides new data that supports global precipitation patterns associated with the Cretaceous Thermal Maximum. This work provides an important framework for much-needed further paleoecological and paleoclimatic analyses into greenhouse conditions in the terrestrial Cretaceous Arctic during this important window in time.