“…The caryopsis consists of pericarp outer layers, seed coat, embryo (germ), and the endosperm (Juliano, 2016; Upadhyay & Karn, 2018). Rice is an important crop used as staple food in many countries, and following dehusking and milling of the paddy rice, the resultant husk (20% of harvest and rich in silica) and bran (10% of harvest) can be used in industries such as agriculture (as biological control agents, soil fertilizers, mushroom subdtrates and mulching), food and feed, water refining, dye removal, fuel and other energy resources, construction materials (ceramics, concrete, and firebrick) (Abdul Wahab et al., 2020; Edrisi et al., 2018; Urtecho et al., 2019; Hossain et al., 2021; Lat Reano, 2020; Sala et al., 2020; Win et al., 2019), and for remediation purposes (Carraro et al., 2021; Pham, 2020; Shamsollahi & Partovinia, 2019; Yulnafatmawita et al., 2020) in a circular bioeconomy fashion. With many available genetic resources and a completely sequenced genome, rice has been considered as a model system for grass cell wall research (Li et al., 2018).…”